Destroyed The Destroyer
|Killed the Vault Boss|
Story related, can't be missed.
When fighting the final boss, aim for the large eye at the top to deal a critical hit with each successive shot. Keep your distance to avoid its tentacle, but be aware that its shockwave cannot be dodged. If you're a hunter, however, a health-regen class mod will help against this. Best to be done with multiple players on the second playthrough.
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